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Lolium --- Lolium --- Lolium perenne --- Lolium perenne --- Puccinia --- Puccinia --- Disease resistance --- Disease resistance --- genetic markers --- genetic markers --- Heritability --- Heritability --- Puccinia coronata --- Puccinia coronata --- Lolium multiflorum --- Lolium multiflorum --- Lolium boucheanum --- Lolium boucheanum